New Data on Enosuchid Anthracosauromorphs (Amphibia) of the Middle–Late Permian of European Russia: Part 2. New Taxa of Enosuchidae

Abstract

New taxa of the family Enosuchidae (Anthracosauromorpha), Enosuchus alveolatus sp. nov. (Sundyr-1 locality, Mari El) and Kitiakia firma gen. et sp. nov. (Bolshoi Kityak locality, Kirov Region), are described based on fragmentary remains from the Middle and Upper Permian of European Russia. The diagnosis of the family Enosuchidae and its stratigraphic distribution are improved; according to the new data, its range is restricted to the Upper Kazanian–Upper Severodvinian (Lower Putyatino). The genus Nyctiboetus Tchudinov, 1955 is excluded from Enosuchidae, since it is considered to be close to gephyrostegid anthracosauromorphs from the Upper Carboniferous of Central Europe.