A new species of the genus Sequoiadendron Buchholz (Cupressaceae) from the Upper Cretaceous of the Enmyvaam River Basin, Central Chukotka

Abstract

A new conifer species, Sequoiadendron tchucoticum A. Sokolova from the Emuneret Formation on the Enmyvaam River (Anadyr River Basin) is described. It is established based on a thorough study of the morphology and epidermal features of shoot impressions, using scanning electron microscopy. The new species has polymorphic shoots with helical branching and three main leaf types: scalelike, subulate, and falcate. Leaves are amphistomatic; amphicyclic stomata are mostly obliquely arranged; ordinary epidermal cells are rectangular, frequently with oblique end walls, and their length-to-width ratio reaches 15: 1. Shoots of the new species prevail among conifers of the Ust’-Emuneret Assemblage. The age of the deposits enclosing the assemblage is debated; according to the latest data, it is estimated as Santonian–Early Campanian.